Zachary Quinto Dr. Oliver Thredson Actor Joseph Fiennes Monsignor Timothy Howard Actor Sarah Paulson Lana Winters Actor Evan Peters Kit Walker Actor Lily Rabe Sister Mary Eunice Actor Lizzie Brocheré Grace Actor James Cromwell Dr. Arthur Arden Actor Jessica Lange Sister Jude Actor Chloë Sevigny Shelley Guest Star Jenna Dewan Teresa Morrison Guest Star Mark Consuelos Spivey Guest Star Mark Margolis Sam Goodman Guest Star Amy Farrington Mrs. Reynolds Guest Star Nikki Hahn Jenny Reynolds Guest Star Bob Bancroft Harry Kirkland Guest Star Eugene Byrd Greyson's Partner Guest Star David Gianopoulos Detective John Greyson Guest Star Kirk Bovill Phil Guest Star Kasey Mahaffy Father James Guest Star Michael Duisenberg Male Uniform Guest Star Ryan Murphy Executive Producer Brad Falchuk Executive Producer Dante Di Loreto Executive Producer Tim Minear Executive Producer