Jon Amiel Director Paul Bettany Charles Darwin Actor Jennifer Connelly Emma Darwin Actor Jeremy Northam Reverend Innes Actor Toby Jones Thomas Huxley Actor Benedict Cumberbatch Joseph Hooker Actor Martha West Annie Darwin Actor Jim Carter Parslow Actor Teresa Churcher Mrs. Davies Actor Harrison Sansostri Lenny Darwin Actor John Collee Screenwriter Jeremy Thomas Producer Peter Watson Executive Producer Christina Yao Executive Producer Janice Eymann Executive Producer Jamie Laurenson Executive Producer David M. Thompson Executive Producer Christopher Young Original Music Jess Hall Cinematographer Melanie Ann Oliver Film Editing Celestia Fox Casting Laurence Dorman Production Design Bill Crutcher Art Director Dominic Capon Set Decoration Louise Stjernsward Costume Design